|c. 30 million|
|Regions with significant populations|
|Bosnia and Herzegovina, Bulgaria, Croatia, Montenegro, North Macedonia, Serbia, Slovenia|
|East South Slavic languages:|
West South Slavic languages:
Serbo-Croatian (Serbian, Croatian, Bosnian, Montenegrin)
(Bulgarians, Macedonians, Montenegrins and Serbs)
(Croats and Slovenes)
(Bosniaks, Pomaks, Gorani and Torbe?i)
|Related ethnic groups|
The South Slavs are a subgroup of Slavic peoples who speak the South Slavic languages. They inhabit a contiguous region in the Balkan Peninsula and the eastern Alps, and in the modern era are geographically separated from the body of West Slavic and East Slavic people by the Romanians, Hungarians, and Austrians in between. The South Slavs today include the nations of Bosniaks, Bulgarians, Croats, Macedonians, Montenegrins, Serbs and Slovenes. They are the main population of the Eastern and Southeastern European countries of Bosnia and Herzegovina, Bulgaria, Croatia, Montenegro, North Macedonia, Serbia and Slovenia.
In the 20th century, the country of Yugoslavia (literally "South Slavia" or "Southern Slavland") united the regions inhabited by South Slavic nations--with the key exception of Bulgaria--into a single state. The concept of Yugoslavia, a single state for all South Slavic peoples, emerged in the late 17th century and gained prominence through the 19th-century Illyrian movement. The Kingdom of Serbs, Croats and Slovenes, renamed the Kingdom of Yugoslavia in 1929, was proclaimed on 1 December 1918, following the unification of the State of Slovenes, Croats and Serbs with the kingdoms of Serbia and Montenegro.
The South Slavs are known in Serbian, Macedonian and Montenegrin as Ju?ni Sloveni (Cyrillic: ?); in Bulgarian as Yuzhni Slavyani (Cyrillic: ? ?); in Croatian and Bosnian as Ju?ni Slaveni; in Slovene as Ju?ni Slovani. The Slavic root *jug? means "south". The Slavic ethnonym itself was used by 6th-century writers to describe the southern group of Early Slavs (the Sclaveni); West Slavs were called Veneti and East Slavs Antes. The South Slavs are also called "Balkan Slavs", although this term does not encompass the Slovenes.
Another name popular in the early modern period was "Illyrians", the name of a pre-Slavic Balkan people, a name first adopted by Dalmatian intellectuals in the late 15th century to refer to South Slavic lands and population. It was then used by the Habsburg Monarchy, France, and notably adopted by the 19th-century Croatian nationalist and Pan-Slavist Illyrian movement. Eventually, the idea of Yugoslavism appeared, aimed at uniting all South Slav-populated territories into a common state. From this idea emerged Yugoslavia, which however did not include Bulgaria.
The Proto-Slavic homeland is the area of Slavic settlement in Central and Eastern Europe during the first millennium AD, with its precise location debated by archaeologists, ethnographers and historians. None of the proposed homelands reaches the Volga River in the east, over the Dinaric Alps in the southwest or the Balkan Mountains in the south, or past Bohemia in the west. Traditionally, scholars put it in the marshes of Ukraine, or alternatively between the Bug and the Dnieper; however, according to F. Curta, the homeland of the southern Slavs mentioned by 6th-century writers was just north of the Lower Danube. Little is known about the Slavs before the 5th century, when they began to spread out in all directions.
Jordanes, Procopius and other late Roman authors provide the probable earliest references to southern Slavs in the second half of the 6th century. Procopius described the Sclaveni and Antes as two barbarian peoples with the same institutions and customs since ancient times, not ruled by a single leader but living under democracy, while Pseudo-Maurice called them a numerous people, undisciplined, unorganized and leaderless, who did not allow enslavement and conquest, and resistant to hardship, bearing all weathers. They were portrayed by Procopius as unusually tall and strong, of dark skin and "reddish" hair (neither blond nor black), leading a primitive life and living in scattered huts, often changing their residence. Procopius said they were henotheistic, believing in the god of lightning (Perun), the ruler of all, to whom they sacrificed cattle. They went into battle on foot, charging straight at their enemy, armed with spears and small shields, but they did not wear armour.
While archaeological evidence for a large-scale migration is lacking, most present-day historians claim that Slavs invaded and settled the Balkans in the 6th and 7th centuries. According to this dominant narrative, up until the late 560s their main activity across the Danube was raiding, though with limited Slavic settlement mainly through Byzantine colonies of foederati. The Danube and Sava frontier was overwhelmed by large-scale Slavic settlement in the late 6th and early 7th century. What is today central Serbia was an important geo-strategical province, through which the Via Militaris crossed. This area was frequently intruded upon by barbarians in the 5th and 6th centuries. From the Danube, the Slavs commenced raiding the Byzantine Empire on an annual basis from the 520s, spreading destruction, taking loot and herds of cattle, seizing prisoners and taking fortresses. Often, the Byzantine Empire was stretched, defending its rich Asian provinces from Arabs, Persians and others. This meant that even numerically small, disorganised early Slavic raids were capable of causing much disruption, but could not capture the larger, fortified cities. The first Slavic raid south of the Danube was recorded by Procopius, who mentions an attack of the Antes, "who dwell close to the Sclaveni", probably in 518. Sclaveni are first mentioned in the context of the military policy on the Danube frontier of Byzantine Emperor Justinian I (r. 527-565). Throughout the century, Slavs raided and plundered deep into the Balkans, from Dalmatia to Greece and Thrace, and were also at times recruited as mercenaries, fighting the Ostrogoths. Justinian seems to have used the strategy of 'divide and conquer', and the Sclaveni and Antes are mentioned as fighting each other. The Antes are last mentioned as anti-Byzantine belligerents in 545, and the Sclaveni continued to raid the Balkans. In 558 the Avars arrived at the Black Sea steppe, and defeated the Antes between the Dnieper and Dniester. The Avars subsequently allied themselves with the Sclaveni, although there was an episode in which the Sclavene Daurentius (fl. 577-579), the first Slavic chieftain recorded by name, dismissed Avar suzerainty and retorted that "Others do not conquer our land, we conquer theirs [...] so it shall always be for us", and had the Avar envoys slain. By the 580s, as the Slav communities on the Danube became larger and more organized, and as the Avars exerted their influence, raids became larger and resulted in permanent settlement. Most scholars consider the period of 581-584 as the beginning of large-scale Slavic settlement in the Balkans. F. Curta points out that evidence of substantial Slavic presence does not appear before the 7th century and remains qualitatively different from the "Slavic culture" found north of the Danube. In the mid-6th century, the Byzantines re-asserted their control of the Danube frontier, thereby reducing the economic value of Slavic raiding. This growing economic isolation, combined with external threats from the Avars and Byzantines, led to political and military mobilisation. Meanwhile, the itinerant form of agriculture (lacking crop rotation) may have encouraged micro-regional mobility. Seventh-century archaeological sites show earlier hamlet collections evolving into larger communities with differentiated zones for public feasts, craftmanship, etc. It has been suggested that the Sclaveni were the ancestors of the Serbo-Croatian group while the Antes were that of the Bulgarian Slavs, with much mixture in the contact zones. The diminished pre-Slavic inhabitants, also including Romanized native peoples,[a] fled from the barbarian invasions and sought refuge inside fortified cities and islands, whilst others fled to remote mountains and forests and adopted a transhumant lifestyle. The Romance speakers within the fortified Dalmatian city-states managed to retain their culture and language for a long time. Meanwhile, the numerous Slavs mixed with and assimilated the descendants of the indigenous population.
Subsequent information about Slavs' interaction with the Greeks and early Slavic states comes from the 10th-century De Administrando Imperio (DAI) by Emperor Constantine VII Porphyrogenitus, the 7th-century compilations of the Miracles of Saint Demetrius (MSD) and the History by Theophylact Simocatta. DAI mentions the beginnings of the Croatian, Serbian and Bulgarian states from the early 7th to the mid-10th century. MSD and Theophylact Simocatta mention the Slavic tribes in Thessaly and Macedonia at the beginning of the 7th century. The 9th-century Royal Frankish Annals (RFA) also mention Slavic tribes in contact with the Franks.
By 700 AD, Slavs had settled in most of Central and Southeast Europe, from Austria even down to the Peloponnese of Greece, and from the Adriatic to the Black Sea, with the exception of the coastal areas and certain mountainous regions of the Greek peninsula. The Avars, who arrived in Europe in the late 550s and had a great impact in the Balkans, had from their base in the Carpathian plain, west of main Slavic settlements, asserted control over Slavic tribes with whom they besieged Roman cities. Their influence in the Balkans however diminished by the early 7th century and they were finally defeated and disappeared as a power at the turn of the 9th century by Bulgaria and the Frankish Empire. The first South Slavic polity and regional power was Bulgaria, a state formed in 681 as a union between the much numerous Slavic tribes and the bulgars of Khan Asparuh. The scattered Slavs in Greece, the Sklavinia, were Hellenized. Romance-speakers lived within the fortified Dalmatian city-states. Traditional historiography, based on DAI, holds that the migration of Serbs and Croats to the Balkans was part of a second Slavic wave, placed during Heraclius' reign.
Inhabiting the territory between the Franks in the north and Byzantium in the south, the Slavs were exposed to competing influences. In 863 to Christianized Great Moravia were sent two Byzantine brothers monks Saints Cyril and Methodius, Slavs from Thessaloniki on missionary work. They created the Glagolitic script and the first Slavic written language, Old Church Slavonic, which they used to translate Biblical works. At the time, the West and South Slavs still spoke a similar language. The script used, Glagolitic, was capable of representing all Slavic sounds, however, it was gradually replaced in Bulgaria in the 9th century, in Russia by the 11th century Glagolitic survived into the 16th century in Croatia, used by Benedictines and Franciscans, but lost importance during the Counter-Reformation when Latin replaced it on the Dalmatian coast. Cyril and Methodius' disciples found refuge in already Christian Bulgaria, where the Old Church Slavonic became the ecclesiastical language.Early Cyrillic alphabet was developed during the 9th century AD at the Preslav Literary School in Bulgaria. The earliest Slavic literary works were composed in Bulgaria, Duklja and Dalmatia. The religious works were almost exclusively translations, from Latin (Croatia, Slovenia) and especially Greek (Bulgaria, Serbia). In the 10th and 11th centuries the Old Church Slavonic leaded to the creation of various regional forms like Serbo-Croatian and Slovenian Economic, religious and political centres of Ohrid and Plovdiv contributed to the important literary production in the Bulgarian Empire. The Bogomil sect, derived from Manichaeism, was deemed heretical, but managed to spread from Bulgaria to Bosnia (where it gained a foothold). and France (Cathars).
Carinthia came under Germanic rule in the 10th century and came permanently under Western (Roman) Christian sphere of influence. What is today Croatia came under Eastern Roman (Byzantine) rule after the Barbarian age, and while most of the territory was Slavicized, a handful of fortified towns, with mixed population, remained under Byzantine authority and continued to use Latin. Dalmatia, now applied to the narrow strip with Byzantine towns, came under the Patriarchate of Constantinople, while the Croatian state remained pagan until Christianization during the reign of Charlemagne, after which religious allegiance was to Rome. Croats threw off Frankish rule in the 9th century and took over the Byzantine Dalmatian towns, after which Hungarian conquest led to Hungarian suzerainty, although retaining an army and institutions. Croatia lost much of Dalmatia to the Republic of Venice which held it until the 18th century. Hungary governed Croatia through a duke, and the coastal towns through a ban. A feudal class emerged in the Croatian hinterland in the late 13th century, among whom were the Kurjakovi?, Ka?i? and most notably the ?ubi?. Dalmatian fortified towns meanwhile maintained autonomy, with a Roman patrician class and Slavic lower class, first under Hungary and then Venice after centuries of struggle.
Ibn al-Faqih described two kinds of South Slavic people, the first of swarthy complexion and dark hair, living near the Adriatic coast, and the other as light, living in the hinterland.
After Ottoman expansion into Byzantine territories in the east in the first half of the 14th century, the internally divided Bulgarian Empire and the short-lived and crumbling Serbian Empire stood next. In 1371, the Ottomans defeated a large Serbian army at the Battle of Maritsa, and in 1389 defeated the Serbian army at the Battle of Kosovo. By now, Serbian and Bulgarian rulers became Ottoman vassals, the southern Serbian provinces and Bulgaria holding out until annexation in the 1390s. The Ottomans conquered Constantinople (1453), Greece (1453-60), the Serbian Despotate (1459) and Bosnia (1463). Much of the Balkans was under Ottoman rule throughout the early modern period. Ottoman rule lasted from the 14th into the early 20th century in some territories. Ottoman society was multi-ethnic and multi-religious, and confessional groups were divided according to the millet system, in which Orthodox Christians (Greeks, Bulgarians, Serbs, etc.) constituted the Rum Millet. In Islamic jurisprudence, the Christians had dhimmi status, which entailed certain taxes and lesser rights. Through Islamization, communities of Slavic Muslims emerged, which survive until today in Bosnia, south Serbia, North Macedonia, and Bulgaria.
In the 16th century, the Habsburg Monarchy controlled what is today Slovenia, Croatia and northern Serbia. The Kingdom of Croatia, which included smaller parts of what is today Croatia, was a crown land of the Habsburg emperor. The early modern period saw large-scale migrations of Orthodox Slavs (mainly Serbs) to the north and west. The Military Frontier was set up as the cordon sanitaire against Ottoman incursions. There were several rebellions against Ottoman rule, but it was not until the 18th century that parts of the Balkans, namely Serbia, were liberated for a longer period. While Pan-Slavism has its origins in the 17th-century Slavic Catholic clergymen in the Republic of Venice and Republic of Ragusa, it crystallized only in the mid-19th century amidst rise of nationalism in the Ottoman and Habsburg empires.
South Slavs are divided linguistically into eastern (Bulgarian and Macedonian) and western groups (Slovenes, Croatian, Bosnian, Serbian, and Montenegrin), and religiously into Orthodox (Serbs, Bulgarians, Macedonians, Montenegrins), Catholic (Croats, Slovenes) and Muslim (Bosniaks, Pomaks, and other minorities). There are an estimated 35 million South Slavs and their descendants living worldwide. Among South Slavic ethnic groups that are also nations are the Serbs, Bulgarians, Croats, Bosniaks, Slovenes, Macedonians and Montenegrins. Bosniaks, Serbs and Croats are the constituent nations of Bosnia and Herzegovina. Among South Slavic minorities or self-identifications are the Yugoslavs (former Yugoslavia), ethnic Muslims (former Yugoslavia), Torbeshi (North Macedonia), Pomaks (Bulgaria, Greece) and Gorani (Kosovo). The Catholic Bunjevci and ?okci, concentrated in northern Serbia and eastern Croatia, are divided between Croat and local identities. There are also smaller communities of West and East Slavic peoples in northern Serbia.
There are seven countries in which South Slavs are the main population:
In addition, there are local South Slavic minorities in non-South Slavic neighbouring countries such as:
Largest cities in South Slavic states
estimates for 1 October 2015
The religious and cultural diversity of the region the South Slavs inhabit has had a considerable influence on their religion. Originally a polytheistic pagan people, the South Slavs have also preserved many of their ancient rituals and traditional folklore, often intermixing and combining it with the religions they later converted to.
Today, the majority of South Slavs are Orthodox Christians; this includes most Bulgarians, Macedonians, Serbs and Montenegrins. Most Slovenes and Croats are Roman Catholics. Bosniaks and other minor ethnic groups (Gorani, ethnic Muslims) and sub-groups (Torbesh and Pomaks) are Muslims. Some South Slavs are atheist, agnostic and/or non-religious.
The South Slavic languages, one of three branches of the Slavic languages family (the other being West Slavic and East Slavic), form a dialect continuum. It comprises, from west to east, the official languages of Slovenia, Croatia, Bosnia and Herzegovina, Montenegro, Serbia, North Macedonia, and Bulgaria. The South Slavic languages are geographically divided from the rest of the Slavic languages by areas where Germanic (Austria), Hungarian and Romanian languages prevail.
South Slavic standard languages are:
The Serbo-Croatian varieties have strong structural unity and are regarded by most linguists as constituting one language. Today, language secessionism has led to the codification of several distinct standards: Serbian, Croatian, Bosnian and Montenegrin. These Serbo-Croatian standards are all based on the Shtokavian dialect group. Other dialect groups, which have lower intelligibility with Shtokavian, are Chakavian in Dalmatia and Kajkavian in Croatia proper. The dominance of Shtokavian across Serbo-Croatian speaking lands is due to historical westward migration during the Ottoman period. Slovene is South Slavic but has many features shared with West Slavic languages. The Prekmurje Slovene and Kajkavian are especially close, and there is no sharp delineation between them. In southeastern Serbia, dialects enter a transitional zone with Bulgarian and Macedonian, with features of both groups, and are commonly called Torlakian. The Eastern South Slavic languages are Bulgarian and Macedonian. Bulgarian has retained more archaic Slavic features in relation to the other languages. Bulgarian has two main yat splits. Slavic Macedonian was codified in Communist Yugoslavia in 1945 and was historically classified as Bulgarian. The Macedonian dialects, divided into three main groups, are regarded overall as being transitional to Bulgarian and Serbo-Croatian. The westernmost Bulgarian dialects (called Shopi) share features with Serbo-Croatian. Furthermore, in Greece there is a notable Slavic-speaking population in Greek Macedonia and western Thrace. Balkan Slavic languages are part of a "Balkan sprachbund" with areal features shared with other non-Slavic languages in the Balkans.
According to the 2013 autosomal IBD survey "of recent genealogical ancestry over the past 3,000 years at a continental scale", the speakers of Serbo-Croatian language share a very high number of common ancestors dated to the migration period approximately 1,500 years ago with Poland and Romania-Bulgaria cluster among others in Eastern Europe. It is concluded to be caused by the Hunnic and Slavic expansion, which was a "relatively small population that expanded over a large geographic area", particularly "the expansion of the Slavic populations into regions of low population density beginning in the sixth century" and that it is "highly coincident with the modern distribution of Slavic languages". According to Kushniarevich et al. 2015, the Hellenthal et al. 2014 IBD analysis also found "multi-directional admixture events among East Europeans (both Slavic and non-Slavic), dated to around 1,000-1,600 YBP" which coincides with "the proposed time-frame for the Slavic expansion". The Slavic influence is "dated to 500-900 CE or a bit later with over 40-50% among Bulgarians, Romanians, and Hungarians". The 2015 IBD analysis found that the South Slavs have lower proximity to Greeks than with East and West Slavs and that there's an "even patterns of IBD sharing among East-West Slavs-'inter-Slavic' populations (Hungarians, Romanians and Gagauz)-and South Slavs, i.e. across an area of assumed historic movements of people including Slavs". The slight peak of shared IBD segments between South and East-West Slavs suggests a shared "Slavonic-time ancestry". The 2014 IBD analysis comparison of Western Balkan and Middle Eastern populations also found negligible gene flow between 16th and 19th century during the Islamization of the Balkans.
According to a 2014 admixture analysis of Western Balkan, the South Slavs show a genetic uniformity. The Bosnians and Croatians were more close to East European populations and largely overlapped with Hungarians from Central Europe. In the 2015 analysis, they formed a western South Slavic cluster with the Bosnians and Slovenians in comparison to eastern cluster formed by Macedonians and Bulgarians with Serbians in the middle. The western cluster has an inclination toward Hungarians, Czechs, and Slovaks, while the eastern cluster toward Romanians and some extent Greeks. The modeled ancestral genetic component of Balto-Slavs among South Slavs was between 55-70%. In the 2018 analysis of Slovenian population, the Slovenian population clustered with Croatians, Hungarians and was close to Czech.
The 2006 Y-DNA study results "suggest that the Slavic expansion started from the territory of present-day Ukraine, thus supporting the hypothesis that places the earliest known homeland of Slavs in the basin of the middle Dnieper". According to genetic studies until 2020, the distribution, variance and frequency of the Y-DNA haplogroups R1a and I2 and their subclades R-M558, R-M458 and I-CTS10228 among South Slavs are in correlation with the spreading of Slavic languages during the medieval Slavic expansion from Eastern Europe, most probably from the territory of present day Ukraine and Southeastern Poland.
The Psalter and the Book of Prophets were adapted or "modernized" with special regard to their use in Bulgarian churches, and it was in this school that glagolitic writing was replaced by the so-called Cyrillic writing, which was more akin to the Greek uncial, simplified matters considerably and is still used by the Orthodox Slavs.
S7.6 "East Europe": The difference between the 'East Europe I' and 'East Europe II' analyses is that the latter analysis included the Polish as a potential donor population. The Polish were included in this analysis to reflect a Slavic language speaking source group." "We speculate that the second event seen in our six Eastern Europe populations between northern European and southern European ancestral sources may correspond to the expansion of Slavic language speaking groups (commonly referred to as the Slavic expansion) across this region at a similar time, perhaps related to displacement caused by the Eurasian steppe invaders (38; 58). Under this scenario, the northerly source in the second event might represent DNA from Slavic-speaking migrants (sampled Slavic-speaking groups are excluded from being donors in the EastEurope I analysis). To test consistency with this, we repainted these populations adding the Polish as a single Slavic-speaking donor group ("East Europe II" analysis; see Note S7.6) and, in doing so, they largely replaced the original North European component (Figure S21), although we note that two nearby populations, Belarus and Lithuania, are equally often inferred as sources in our original analysis (Table S12). Outside these six populations, an admixture event at the same time (910CE, 95% CI:720-1140CE) is seen in the southerly neighboring Greeks, between sources represented by multiple neighboring Mediterranean peoples (63%) and the Polish (37%), suggesting a strong and early impact of the Slavic expansions in Greece, a subject of recent debate (37). These shared signals we find across East European groups could explain a recent observation of an excess of IBD sharing among similar groups, including Greece, that was dated to a wide range between 1,000 and 2,000 years ago (37)
However, a study by Battaglia et al. (2009) showed a variance peak for I2a1 in the Ukraine and, based on the observed pattern of variation, it could be suggested that at least part of the I2a1 haplogroup could have arrived in the Balkans and Slovenia with the Slavic migrations from a homeland in present-day Ukraine... The calculated age of this specific haplogroup together with the variation peak detected in the suggested Slavic homeland could represent a signal of Slavic migration arising from medieval Slavic expansions. However, the strong genetic barrier around the area of Bosnia and Herzegovina, associated with the high frequency of the I2a1b-M423 haplogroup, could also be a consequence of a Paleolithic genetic signal of a Balkan refuge area, followed by mixing with a medieval Slavic signal from modern-day Ukraine.
R1a-M458 exceeds 20% in the Czech Republic, Slovakia, Poland, and Western Belarus. The lineage averages 11-15% across Russia and Ukraine and occurs at 7% or less elsewhere (Figure 2d). Unlike hg R1a-M458, the R1a-M558 clade is also common in the Volga-Uralic populations. R1a-M558 occurs at 10-33% in parts of Russia, exceeds 26% in Poland and Western Belarus, and varies between 10 and 23% in the Ukraine, whereas it drops 10-fold lower in Western Europe. In general, both R1a-M458 and R1a-M558 occur at low but informative frequencies in Balkan populations with known Slavonic heritage.
Hg I2a1a2b-L621 was present in 5 Conqueror samples, and a 6th sample form Magyarhomorog (MH/9) most likely also belongs here, as MH/9 is a likely kin of MH/16 (see below). This Hg of European origin is most prominent in the Balkans and Eastern Europe, especially among Slavic speaking groups.
Az I2-CTS10228 (köznevén ,,dinári-kárpáti") alcsoport legkorábbi közös ?se 2200 évvel ezel?ttre tehet?, így esetében nem arról van szó, hogy a mezolit népesség Kelet-Európában ilyen mértékben fennmaradt volna, hanem arról, hogy egy, a mezolit csoportoktól származó sz?k család az európai vaskorban sikeresen integrálódott egy olyan társadalomba, amely hamarosan er?teljes demográfiai expanzióba kezdett. Ez is mutatja, hogy nem feltétlenül népek, mintsem családok sikerével, nemzetségek elterjedésével is számolnunk kell, és ezt a jelenlegi etnikai identitással összefüggésbe hozni lehetetlen. A csoport elterjedése alapján valószín?síthet?, hogy a szláv népek migrációjában vett részt, így válva az R1a-t követ?en a második legdominánsabb csoporttá a mai Kelet-Európában. Nyugat-Európából viszont teljes mértékben hiányzik, kivéve a kora középkorban szláv nyelvet beszél? keletnémet területeket.
Based on SNP analysis, the CTS10228 group is 2200 ± 300 years old. The group's demographic expansion may have begun in Southeast Poland around that time, as carriers of the oldest subgroup are found there today. The group cannot solely be tied to the Slavs, because the proto-Slavic period was later, around 300-500 CE... The SNP-based age of the Eastern European CTS10228 branch is 2200 ± 300 years old. The carriers of the most ancient subgroup live in Southeast Poland, and it is likely that the rapid demographic expansion which brought the marker to other regions in Europe began there. The largest demographic explosion occurred in the Balkans, where the subgroup is dominant in 50.5% of Croatians, 30.1% of Serbs, 31.4% of Montenegrins, and in about 20% of Albanians and Greeks. As a result, this subgroup is often called Dinaric. It is interesting that while it is dominant among modern Balkan peoples, this subgroup has not been present yet during the Roman period, as it is almost absent in Italy as well (see Online Resource 5; ESM_5).
The geographic distributions of the major eastern European NRY haplogroups (R1a-Z282, I2a-P37) overlap with the area occupied by the present-day Slavs to a great extent, and it might be tempting to consider both haplogroups as Slavic-specic patrilineal lineages